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All samples were serially diluted in assay buffer in duplicate and incubated with cells for 30 min at RT before detection with the cAMP dynamic 2 HTRF assay (Cisbio assays, Bedford, MA) as described previously 51

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Most strikingly, Y391 GH5 formed a tight hydrogen bond with a water molecule, which connects to E281 3.50b and H214 2.50b in the GLP-2RG s complex (Supplementary information, Fig