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Loren Pickart, a biochemist, first isolated glycyl-l-histidyl-l-lysine (GHK) from the plasma in 1973 [29]

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However, other studies suggest that cyclosporin A upregulates gingival SHH (Sonic hedgehog protein) expression in vitro, while in vivo that inhibition of the SHH pathway counteracts the stimulatory effect of cyclosporin A on gingival fibroblast proliferation

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Shanmuganathan, D., & Sivakumaran, N